PUBLICATIONS

1. Huang X, Platin R, Unger A, Shenkar N, Zhan A (2025). Interplay of multilayered transcriptomic plasticity in response to temperature fluctuations during biological invasions: A case study in Invasive Styela plicata. Mol Ecol e70013.‏
2. Lee SSC, Unger A, Teo SLM, Shenkar N (2025) Culturing the solitary ascidian Phallusia nigra in closed and open water systems for tropical environmental research. L & O Methods  https://doi.org/10.1002/lom3.10713
3. Berenshtein I, Shenkar N (2025) Modeling non-indigenous ascidian larval dispersal in the coastal waters of the Eastern Mediterranean Sea. Diversity and Distributions https://doi.org/10.1111/ddi.70009
4. Golanski D, Nachmias A, Kahn G, Fireman A, Hepner Ucko O, Shenkar N, Yahel G (2025) Massive colonization by the solitary ascidian Microcosmus exasperatus Heller, 1878, on the sandy bottom of the Israeli littoral. In: Fowler A, Robinson T, Bortolus A, Canning-Clode J, Therriault T (Eds) Proceedings of the 11th International Conference on Marine Bioinvasions. Aquatic Invasions 20(1): 53-68. https://doi.org/10.3391/ai.2025.20.1.131343
5. Harel E, Zucker I, Shenkar N (2024) Effects of biological filtration by ascidians on microplastic composition in the water column. Chemosphere 367 143589 
6. Vered G, Nordland O, Gozlan I, Shenkar N (2024) Occurrence of plastic additives in coral-reef invertebrates on natural and plastic substrates.
   Mar Pollut Bull 208: 116935
7. Navon G, Nordland O, Kaplan A, Avisar D, Shenkar N (2024) Detection of 10 commonly used pharmaceuticals in reef-building stony corals from shallow (5-12 m) and deep (30-40 m) sites in the Red Sea. Environ Pollut 124698
8. Montesanto F, Gissi C, Mastrototaro F, Hadjez R, Huchon D, Novak L, Shenkar N (2023) Against the flow: Integrated taxonomy and environmental suitability of a cold-water species dispersing to warmer waters. Biol Inv 1-23 https://doi.org/10.1007/s10530-023-03222-0 
9. Platin R, Shenkar N (2023) Can stand the heat–ecology of the potentially invasive ascidian Styela plicata along the Mediterranean coast of Israel. Front Mar Sci 10: 1159231
10. Galanidi M et al. (+30 co-authors) (2023) Validated inventories of Non-Indigenous Species (NIS) for the Mediterranean Sea as tools for regional policy and patterns of NIS spread. Diversity 15, 962. https://doi.org/10.3390/d15090962
11. Shenkar N (2023) Sea squirts adventures. Genesis e23560‏
12. ​Vered G, Shenkar N (2023) Plastic pollution in a coral reef climate refuge: Occurrence of anthropogenic debris, microplastics, and plasticizers in the Gulf of Aqaba. Sci Tot Environ 16779
13. Bereza D, Grey E, Shenkar N (2023) Prioritizing management of high-risk routes and ports by vessel type to improve marine biosecurity efforts. J Environ Manage 337 117597
14. Huang X, Li H, Shenkar N, Zhan A (2023) Multidimensional plasticity jointly contributes to rapid acclimation to environmental challenges during biological invasions. RNA doi 10.1261/rna.079319.122
15. Gordon T, Zaquin T, Kowarsky MA, Voskoboynik Y, Hendin N, Wurtzel O, Caicci F, Manni L, Voskoboynik A, Manni L, Shenkar N (2022) Stemness activity underlying whole brain regeneration in a basal chordate. Cells 11(23), 3727
16. Vered G, Shenkar N (2022) Limited effects of environmentally-relevant concentrations in seawater of dibutyl phthalate, dimethyl phthalate, bisphenol A, and 4-nonylphenol on the reproductive products of coral-reef organisms. Environ Pollut 314: 120285
17. Hendin N, Gordon T, Shenkar N, Wurtzel O (2022) Molecular characterization of the immediate wound response of the solitary ascidian Polycarpa mytiligera Dev Dyn 2022 1-14
18. Bereza D, Shenkar N (2022)  Shipping voyage simulation reveals abiotic barriers to marine bioinvasions. Sci Tot Environ 837: 155741‏
19. Shenkar N, Bereza D, Gordon T, Koplovitz G, Navon G, Novak L, Vered G (2022) Ascidians of the Red Sea: in peril and invasive. In Imperiled the Encyclopedia of Conservation. Elsevier https://doi.org/10.1016/B978-0-12-821139-7.00232-4​
20. Vered G, Shenkar N (2021) Monitoring plastic pollution in the oceans. Curr Opin Toxicol 27:60-68
21. Navon G, Novak L, Shenkar N (2021). Proteomic changes in the solitary ascidian Herdmania momus following exposure to the anticonvulsant medication carbamazepine. Aquat Toxicol 237: 105886
22. Gordon T, Upadhyay AK, Manni L, Huchon D, Shenkar N (2021) And then there were three…: Extreme regeneration ability of the solitary chordate Polycarpa mytiligera Front Cell Dev Biol 9: 793‏
23. Ben-Tal A, Shenkar N, Paz A, Conley K, Sutherland K, Yahel G (2021) High mucous-mesh production by the ascidian Herdmania momus. Mar Ecol Prog Ser 663:223-228
24. Jacobi Y, Shenkar N, Ward EJ, Rosa M, Ramon GZ, Shavit U, Yahel G (2021) Evasive plankton: Size‐independent particle capture by ascidians. Limnol and Oceanogr https://doi.org/10.1002/lno.11658
25. Anderson G, Shenkar N (2021) Potential effects of biodegradable single-use items in the sea: polylactic acid (PLA) and solitary ascidians. Environ Pollut 268: 115364‏
26. Navon G, Kaplan A, Avisar D, Shenkar N (2020) Assessing pharmaceutical contamination along the Mediterranean and Red Sea coasts of Israel: Ascidians (Chordata, Ascidiacea) as bioindicators Mar Pollut Bull Article 160: 111510 
27. Bereza D, Rosen D, Shenkar N (2020) Current trends in ship movement via the Suez Canal in relation to future legislation and mitigation of marine species introductions. Manag Biol Inv 11" 476-492
28. Novak L, Shenkar N (2020) Occurrence of Didemnum perlucidum Monniot F., 1983 on artificial substrates along the Mediterranean coast of Israel. Med Mar Sci http://dx.doi.org/10.12681/mms.22223
29. Gordon T, Roth L, Caicci F, Manny L, Shenkar N  (2020) Spawning induction, development and culturing of the solitary ascidian Polycarpa mytiligera, an emerging model for regeneration studies. Front Zool 17 1-14
30. Tsiamis et al. (2020) Prioritizing marine invasive alien species in the European Union through horizon scanning. Aquat Conserv DOI: 10.1002/aqc.3267
31. Tzafriri-Milo R, Benaltabet T, Torfstein A,  Shenkar N (2019) The potential use of invasive ascidians for biomonitoring heavy-metal pollution. Front Mar Sci doi: 10.3389/fmars.2019.00611
32. Avisar D, Kaplan A,Ronen-Eliraz G, Vered G, Shenkar N, Gozlan I (2019) Validated method for the detection of three phthalates derived from marine invertebrates. Am J Anal Chem 10:445-458
33. Levitt-Barmats Y, Yanai Z, Magory-Cohen T, Shenkar N (2019) Life-history traits and ecological characteristics of the ornamental shrimp Neocaridina denticulata (De Haan, 1844), recently introduced into the freshwater systems of Israel. Aquat Inv 14: 684-702
34. Gilbert P, Porter SM, Sun CY, Xiao S, Gibson BM, Shenkar N, Knoll AH (2019) Biomineralization by particle attachment in early animals. Proc. Natl. Acad. Sci. USA https://doi.org/10.1073/pnas.1902273116  
35. Gordon T, Manny L, Shenkar N (2019) Regeneration ability in four stolidobranch ascidians: Ecological and evolutionary implications. J Exp Mar Biol Ecol 519 151184 https://doi.org/10.1016/j.jembe.2019.151184 
36. Kuplik Z, Novak L, Shenkar N (2019) Proteomic profiling of ascidians as a tool for biomonitoring marine environments. PLoS One 14(4): e0215005.
37. Vered G, Kaplan A, Avisar D, Shenkar N (2019) Using solitary ascidians to assess microplastic and phthalate plasticizers pollution among marine biota: A case study of the Eastern Mediterranean and Red Sea. Mar Pollut Bull 138: 618-625  
38. Chen Y, Shenkar N, Ni P, Lin Y, Li S, Zhan A (2018) Rapid microevolution during recent range expansion to harsh environments. BMC Evol Biol 18: 187
39. Dror H, Novak L, Evans JS, López-Legentil S, Shenkar N (2018) Core and dynamic microbial communities of two invasive ascidians on a wide geographic scale. Microbial Ecology https://doi.org/10.1007/s00248-018-1276-z​
40. Gewing M, Goldstein E, Buba Y, Shenkar N (2018) Temperature resilience facilitates invasion success of the solitary ascidian Herdmania momus. Biol Inv https://doi.org/10.1007/s10530-018-1827-8
41. Shenkar N, Rosen D (2018) How the invention of the shipping container influenced marine bioinvasion. Manag Biol Invasion 9:187-194 https://doi.org/10.3391/mbi.2018.9.3.02
42. Evans JS, Erwin PM, Shenkar N, Lopez-Legentil S (2018) A comparison of symbiotic microbial communities in nonnative and native ascidians from reef and harbor habitats. FEMS Microbiol Ecol doi: 10.1093/femsec/fiy139
43. *Levitt-Barmats Y, Shenkar N* (2018) Investigating the symbiotic relationship between the caridean shrimp Odontonia sibogae (Bruce, 1972) and its ascidian host Herdmania momus (Savigny, 1816). PLoS One 13(2): e0192045
44. Karachle PK, Corsini Foka M, Crocetta F, Dulčić J, Dzhembekova N, Galanidi M, Ivanova P, Shenkar N, Skolka M, Stefanova E, Stefanova K, Surgiu V, Uysal I, Verlaque M, Zenetos A (2017) Setting-up a billboard of marine invasive species in the ESENIAS area: current situation and future expectancies. Acta Adriat 58(3): 429 – 458
45. *Jacobi Y, Yahel G, Shenkar N (2017) Efficient filtration of micron and submicron particles by ascidians from oligotrophic waters. Limnol and Oceanogr doi:10.1002/lno.10736
46. Gewing M, López-Legentil S, Shenkar N* (2017) Anthropogenic factors influencing invasive ascidian establishment in natural environments.  Mar Environ Res 131: 236-242
47. Evans JS, Erwin PM, Shenkar N, López-Legentil S (2017) Introduced ascidians harbor highly diverse and host-specific symbiotic microbial assemblages.  Sci Rep. 7(1):11033
48. Granot I, Shenkar N, Belmaker J (2017) Habitat niche breadth predicts invasiveness in ascidians. Ecol Evol 7:7838–7847
49. Villalobos SM, Lambert G, Shenkar N, López-Legentil S (2017) Distribution and population dynamics of key ascidians in North Carolina harbors and marinas Aquat Inv 12 (4):447-458
50. Shlesinger T, Shenkar N, Loya Y* (2017) Gamete spawning of the ascidian Phallusia nigra in the Red Sea. Bull Mar Sci. Available on line
51. Gewing M, Shenkar N* (2017) Monitoring the magnitude of marine vessel infestation by non-indigenous ascidians in the Mediterranean. Mar Pollut Bull 121:52-59
52. Shmuel Y, Shenkar N* (2017) Reproductive cycle and ecology of the tropical ascidian Halocynthia spinosa in the Red Sea. Mar Biol 164:147
53. Novak L, López-Legentil S, Shenkar N* (2017) Rapid establishment of the non-indigenous ascidian Styela plicata and its associated bacteria in marinas and fishing harbors along the Mediterranean coast of Israel. Mediterr Mar Sci 18: 324-331
54. Zenetos A, Çinar ME, Crocetta F, Golani D, Rosso A, Servello G, Shenkar N, Turon X, Verlaque M (2017) Uncertainties and validation of alien species catalogues: the Mediterranean as an example. Estuarine Coastal and Shelf 191:171-187 
55.  Shenkar N*, Shmuel Y, Huchon D (2017) The invasive ascidian Ciona robusta recorded from a Red Sea marina. Mar Biodivers doi:10.1007/s12526-017-0699-y
56. Tassia MG, Cannon JT, Konikoff CE, Shenkar N, Halanych KM, Swalla BJ* (2016) The Global Diversity of Hemichordata. PLoS ONE 11(10): e0162564. doi:10.1371/journal.pone.0162564
57. Tsiamis* et al. (2016) The EASIN Editorial Board: quality assurance, exchange and sharing of alien species information in Europe. Management of Biological Invasions 7:321-328  
58. Shenkar N*, Koplovitz G, Dray L, Gissi C, Huchon D (2016) Back to solitude- Solving the phylogenetic position of Diazonidae a molecular and developmental approach. Mol Phylo Evol 100: 51-56 
59. Raijman-Nagar L, Shenkar N* (2016) Temperature and salinity sensitivity of the invasive ascidian Microcosmus exasperatus. Aquat Invasions 11(1): 33–43  
60. Raijman-Nagar L, Shenkar N* (2016) From tropical to sub-tropical: prolonged reproductive activiy of the invasive ascidian Microcosmus exasperatus in the eastern Mediterranean.  Frontiers in Ecology and Evolution, http://dx.doi.org/10.3389/fevo.2016.00102
61. Gewing M, Bronstein O, Raijman Nagar L, Granot I, Frid O, Shenkar N* (2016) First record of the non-indigenous ascidian Microcosmus exasperatus, Heller 1878, in Cyprus. Mar Biodivers 46: 937-941
62. Koplovitz G, Shmuel Y, Shenkar N* (2016) Floating docks in tropical environments- a reservoir for the opportunistic ascidian Herdmania momus.  Manag Biol Invasion 7(1): 43-50
63. Shenkar N*, Gordon T (2015) Gut-spilling in chordates: Evisceration in the tropical ascidian Polycarpa mytiligera. Sci Rep doi:10.1038/srep09614 
64. Koplovitz G, Hirose E, Hirose M, Shenkar N* (2015) Being Green in the Red Sea - Ecology of the Photosymbiotic Ascidian Diplosoma simile (Ascidiacea: Didemnidae) in the Gulf of Aqaba. Syst Biodivers 13(2): 131-139
65. Levitt Y*, De Grave S, Shenkar N (2014) First record of an invasive shrimp from the family Processidae (Crustacea, Decapoda) in the Mediterranean Sea. Medit Mar Sci 15: 650-653 
66. Gewing M, Rhotman SBS, Raijman Nagar L, Shenkar N* (2014) Early stages of establishment of the non-indigenous ascidian Herdmania momus (Savigny, 1816) in shallow and deep water environments on natural substrates in the Mediterranean Sea. BioInv Rec 3: 77-81 
67. Rubinstein N, Feldstein T, Shenkar N, Botero Castro F, Griggio F, Mastrototaro F, Delsuc F, Douzery EJP, Gissi C, Huchon D (2013) Deep sequencing of mixed total DNA without barcodes allows efficient assembly of highly plastic ascidian mitochondrial genomes. Genome Biol Evol 5:1185-1199 
68. Shenkar N *(2013) A new species of the genus Rhopalaea (Ascidiacea) from the Red Sea. Zootaxa, 3599 (1): 051–058 
69. Appeltans et al. (+115 co-authors) (2012) Magnitude of global marine biodiversity: one third of sea creatures discovered. Curr Biol 22:1-14 Highlighted in “Nature Research Highlights” Nature 491: 498 
70. Rius M, Shenkar N (2012) Ascidians introduction through the Suez Canal: The case study of an Indo-Pacific species. Mar Pollut Bull 64: 2060-2068 
71. Shenkar N* (2012) Ascidian (Chordata, Ascidiacea) diversity in the Red Sea. Mar Biodiv doi 10.1007/s12526-012-0124-5 
72. Shenkar N*, Swalla BJS (2011) Global diversity of Ascidiacea. PLoS One, 6(6): e20657 
73. Lambert G, Shenkar N, Swalla BJS (2010) First Pacific record of the north Atlantic ascidian Molgula citrina- bioinvasion or circumpolar distribution? Aquatic Inv 5:369-378 
74. Shenkar N, Lambert G (2010) A new species of the genus Boltenia (Ascidiacea) from the Red Sea. Zootaxa 2391: 61–68 
75.  Shenkar N, Swalla BJS (2010) Molecular data confirms synonym in Roscovite molgulid ascidians. Cah Biol Mar 51:85-87 
76. Tsagkogeorga G, Turon X, Hopcroft RR, Tilak M, Feldstein T, Shenkar N, Loya Y, Huchon D, Douzery EJP, Delsuc F (2009) An updated 18S rRNA phylogeny of tunicates based on mixture and secondary structure models. BMC Evol Biol 9:187 
77. Singh TR, Tsagkogeorga G, Delsuc F, Blanquart S, Shenkar  N, Loya Y, Douzery EJP, Huchon D (2009) Tunicate mitogenomics and phylogenetics: peculiarities of the Herdmania momus mitochondrial genome and support for the new chordate phylogeny. BMC Genomics 10:534 
78. Shenkar N*, Loya Y (2009) Non-indigenous ascidians along the Mediterranean coast of Israel. Mar Biodiver Rec 2:1-7 
79. Shenkar N*, Loya Y (2008) The solitary ascidian Herdmania momus: native (Red Sea) vs. non-indigenous (Mediterranean) populations. Biol Inv 10:1431-1439 
80. Shenkar N*, Bronstein O, Loya Y (2008) Population dynamics of a coral reef ascidian in a deteriorating environment. Mar Ecol Prog Ser 367:163-171 
81. Shenkar N*, Zeldman Y, Loya Y (2008) Ascidian recruitment patterns on an artificial reef in Eilat (Red Sea). Biofouling 24:119-128 
82. Shenkar N*, Monniot F (2006) A new species of the genus Botryllus (Ascidiacea) from the Red Sea. Zootaxa 1256:11-19 
83. Shenkar N*, Fine M, Kramarsky-Winter E, Loya Y (2005) Population dynamics of zooxanthellae during a bacterial bleaching event. Coral Reefs 25:223-227 
84. Shenkar N*, Fine M, Loya Y (2005) Size matters: bleaching dynamics of the coral Oculina patagonica. Mar Ecol Prog Ser 294:181-188 

CHAPTERS IN BOOKS
1. Shenkar N, Loya Y (2008) Ecology and systematics of the ascidian fauna in the Gulf of Eilat (Aqaba). In “Aqaba-Eilat, the Improbable Gulf. Environment, Biodiversity and Preservation” Editor FD Por, Magnes, Jerusalem. pp 197-208
2. *Gordon T, Shenkar N (2018) Ascidians as model organisms for regenerative studies. In “Marine Organisms as Model Systems in Biology and Medicine” Editors M Kloc, JZ Kubiak, Springer, Cham. p. 321-336